Tóth AG, Bende A, Hornok S, Wagenhoffer Z, Szulyovszky B, Galla V, Vöröskői P, Keve G. Reports of Lipoptena fortisetosa on dogs and in the environment, and evidence of its widespread establishment in Hungary. Int J Parasitol Parasites Wildl. 2025 Nov 13;28:101161. doi: 10.1016/j.ijppaw.2025.101161.
Szekeres, Sándor, Nagy, Gábor, Csivincsik, Ágnes, Takács, Nóra, Kontschán, Jenő, Hornok, Sándor, The European Badger (Meles meles) as a Host for Ticks and Tick-Borne Pathogens in Peri-Urban Environments, Hungary, Transboundary and Emerging Diseases, 2025, 1587951, 10 pages, 2025. https://doi.org/10.1155/tbed/1587951
Kürtös G, Rácz N, Kassay V, Olaszy K, Vizi Z, Szilasi A, Takács N, Keve G, Tuska-Szalay B, Hornok S. Infection caused by a parasite most closely related to Hepatozoon luiperdjie in cats from Hungary. Ticks Tick Borne Dis. 2025 Nov;16(6):102556. doi: 10.1016/j.ttbdis.2025.102556.
Nagy E, Nagy R R, Miklós M, Szekeres S, Bawan M A, Földvári G, Rózsa L, Fok É, Sréter T, Tari T, Kovács M, Ágnes Csivincsik, Gábor Nagy,
Eye to eye with Thelazia-infected canids in Central European forests,
Current Research in Parasitology & Vector-Borne Diseases,
Volume 9,2026,10.1016/j.crpvbd.2026.100353 https://www.sciencedirect.com/science/article/pii/S2667114X26000099
Hornok S, Kontschán J, Keve G, Takács N, Van Nguyen D, Ho KNP, Görföl T, Wang Y, Farkas R, Dao TTH. First report of Haemaphysalis bispinosa, molecular-geographic relationships of Ixodes granulatus and a new Dermacentor species from Vietnam. Parasit Vectors. 2025 Jan 23;18(1):21. doi: 10.1186/s13071-024-06641-7.
Balázs B, Boros Á, Pankovics P, Nagy G, Szekeres S, Urbán P, Reuter G. Detection and complete genome characterization of a genogroup X (GX) sapovirus (family Caliciviridae) from a golden jackal (Canis aureus) in Hungary. Arch Virol. 2024 Apr 17;169(5):100. doi: 10.1007/s00705-024-06034-2.
Deak G, Ionică AM, Péter Á, Sándor AD, Matei IA, D'Amico G, Liénard E, Gherman CM, Mihalca AD, Bouhsira E. Fleas of wild carnivores in Romania: diversity, distribution, and host-associations. Parasit Vectors. 2024 Mar 21;17(1):148. doi: 10.1186/s13071-024-06235-3.
Dao TTH, Takács N, Tran TN, Truong AN, Skinner K, Kontschán J, Farkas R, Hornok S. Detection of tick- borne pathogens in the pangolin tick, Amblyomma javanense, from Vietnam and Laos, including a novel species of Trypanosoma. Acta Trop. 2024 Dec;260:107384. doi: 10.1016/j.actatropica.2024.107384. Epub 2024 Sep 10.
Cui N, Su L, Wang Z, Hornok S, Tang L, Yang M, Zhang Y, Zhao G, Wang Y. First identification of Cytauxzoon manul in Eurasian lynx (Lynx lynx) in northwestern China. Parasit Vectors. 2024 Jun 6;17(1):249. doi: 10.1186/s13071-024-06326-1.
Liu G, Tan W, Wang H, Han X, Hornok S, Zhao S, Mi L, Wang S, Yang M, Wang Y. (2024) The great gerbil (Rhombomys opimus) as a host for tick species in Gurbantunggut Desert. Parasites and Vectors 17: 1 Paper: 55, 7 p.
https://parasitesandvectors.biomedcentral.com/articles/10.1186/s13071-024-06160-5
Fedorov D, Hornok S. (2024) Checklist of hosts, illustrated geographical range, and ecology of tick species from the genus Ixodes (Acari, Ixodidae) in Russia and other post-Soviet countries.
Zookeys 1201: pp. 255-343., 89 p. https://zookeys.pensoft.net/article/115467/
Tuska-Szalay B, Boldogh SA, Farkas R, Rompos L, Takács N, Beresnyák V, Hornok S (2023). Screening of Domestic Cats from North-Eastern Hungary for Hepatozoon felis and Cytauxzoon europaeus That Cause Infections in Local Wildcat Populations. Pathogens 12(5): 656. http://doi.org/10.3390/pathogens12050656
Hornok S, Kontschán J, Takács N, Heyne H, Kovács ÁB, Plantard O, Halajian A et al. (2023). Molecular-phylogenetic analyses of Ixodes species from South Africa suggest an African origin of bird-associated exophilic ticks (subgenus Trichotoixodes). Parasites and Vectors 16(1). http://doi.org/10.1186/s13071-023-05998-5
Balmos OM, Supeanu Tamba, P, Horvath C, Panait LC, Sandor AD, Cazan CD, Ungur A, Motiu M, Manita FA, Ancuceanu BC, Barbuceanu F, Dhollander S, Mihalca, AD. African Swine Fever Virus Load in Hematophagous Dipterans Collected in Outbreaks from Romania: Risk Factors and Implications. Transbound and Emer. 2023 Feb Paper: 3548109, 9 p. doi: 10.1155/2023/3548109
Hornok S, Mihalca AD, Kontschán J, Takács N, Fedorov D, Plantard O, Sándor AD. Phylogenetic analyses of Ixodes rugicollis with notes on its morphology in comparison with Ixodes cornutus. Parasit Vectors. 2023 Mar 16;16(1):106. doi: 10.1186/s13071-023-05718-z.
Hornok S, Boldogh SA, Takács N, Kontschán J, Szekeres S, Sós E, Sándor AD, Wang Y, Tuska-Szalay B. Molecular epidemiological study on ticks and tick-borne protozoan parasites (Apicomplexa: Cytauxzoon and Hepatozoon spp.) from wild cats (Felis silvestris), Mustelidae and red squirrels (Sciurus vulgaris) in central Europe, Hungary. Parasit Vectors. 2022 May 21;15(1):174. doi: 10.1186/s13071-022-05271-1.
Hornok S, Boldogh SA, Takács N, Sándor AD, Tuska-Szalay B. Zoonotic ecotype-I of Anaplasma phagocytophilum in sympatric wildcat, pine marten and red squirrel - Short communication. Acta Vet Hung. 2022 Aug 26. doi: 10.1556/004.2022.00021.
Hornok S, Meyer-Kayser E, Kontschán J, Takács N, Plantard O, Cullen S, Gaughran A, Szekeres S, Majoros G, Beck R, Boldogh SA, Horváth G, Kutasi C, Sándor AD. Morphology of Pholeoixodes species associated with carnivores in the western Palearctic: Pictorial key based on molecularly identified Ixodes (Ph.) canisuga, I. (Ph.) hexagonus and I. (Ph.) kaiseri males, nymphs and larvae. Ticks Tick Borne Dis. 2021 Jul;12(4):101715. doi: 10.1016/j.ttbdis.2021.101715.
Sang C, Yang Y, Dong Q, Xu B, Liu G, Hornok S, Liu Z, Wang Y, Hazihan W. Molecular survey of Babesia spp. in red foxes (Vulpes Vulpes), Asian badgers (Meles leucurus) and their ticks in China. Ticks Tick Borne Dis. 2021 Jul;12(4):101710. doi: 10.1016/j.ttbdis.2021.101710.
Liu G, Zhao S, Tan W, Hornok S, Yuan W, Mi L, Wang S, Liu Z, Zhang Y, Hazihan W, Gu X, Wang Y. Rickettsiae in red fox (Vulpes vulpes), marbled polecat (Vormela peregusna) and their ticks in northwestern China. Parasit Vectors. 2021 Apr 19;14(1):204. doi: 10.1186/s13071-021-04718-1.
Liu X, Yang M, Song S, Liu G, Zhao S, Liu G, Hornok S, Wang Y, Jiang H. Brucella melitensis in Asian Badgers, Northwestern China. Emerg Infect Dis. 2020 Apr;26(4):804-806. doi: 10.3201/eid2604.190833.
Yan B, Zhu Q, Xu J, Zhao S, Piao D, Chen C, Liu Q, Hornok S, Wang Y, Wang B, Jiang H. Brucella in Himalayan Marmots (Marmota himalayana). J Wildl Dis. 2020 Jul;56(3):730-732. doi: 10.7589/2019-09-237.
Hornok S, Sándor AD, Földvári G, Ionică AM, Silaghi C, Takács N, Schötta AM, Wijnveld M. First broad-range molecular screening of tick-borne pathogens in Ixodes (Pholeoixodes) kaiseri, with special emphasis on piroplasms. Acta Vet Hung. 2020 May 8;68(1):30-33. doi: 10.1556/004.2020.00003.
Hornok S, Mühldorfer K, Takács N, Hofmann-Lehmann R, Meli ML, Gyuranecz M, Unnsteinsdóttir ER, Greenwood AD, Czirják GÁ. Broad Range Screening of Vector-Borne Pathogens in Arctic Foxes (Vulpes lagopus) in Iceland. Animals (Basel). 2020 Nov 4;10(11):2031. doi: 10.3390/ani10112031.
Szekeres S, Docters van Leeuwen A, Tóth E, Majoros G, Sprong H, Földvári G. Road-killed mammals provide insight into tick-borne bacterial pathogen communities within urban habitats. Transbound Emerg Dis. 2019 Jan;66(1):277-286. doi: 10.1111/tbed.13019.
Koetsveld J, Platonov AE, Kuleshov K, Wagemakers A, Hoornstra D, Ang W, Szekeres S, van Duijvendijk GLA, Fikrig E, Embers ME, Sprong H, Hovius JW. Borrelia miyamotoi infection leads to cross-reactive antibodies to the C6 peptide in mice and men. Clin Microbiol Infect. 2020 Apr;26(4):513.e1-513.e6. doi: 10.1016/j.cmi.2019.07.026.
Hornok S, Horváth G, Takács N, Kontschán J, Szőke K, Farkas R. Molecular identification of badger-associated Babesia sp. DNA in dogs: updated phylogeny of piroplasms infecting Caniformia. Parasit Vectors. 2018 Apr 11;11(1):235. doi: 10.1186/s13071-018-2794-8.
Hornok S, Horváth G, Takács N, Farkas R, Szőke K, Kontschán J. Molecular evidence of a badger-associated Ehrlichia sp., a Candidatus Neoehrlichia lotoris-like genotype and Anaplasma marginale in dogs. Ticks Tick Borne Dis. 2018 Jul;9(5):1302-1309. doi: 10.1016/j.ttbdis.2018.05.012.
Liu X, Yang M, Liu G, Zhao S, Yuan W, Xiao R, Hazihan W, Hornok S, Wang Y. Molecular evidence of Rickettsia raoultii, "Candidatus Rickettsia barbariae" and a novel Babesia genotype in marbled polecats (Vormela peregusna) at the China-Kazakhstan border. Parasit Vectors. 2018 Aug 4;11(1):450. doi: 10.1186/s13071-018-3033-z.
Foley P, Foley J, Sándor AD, Ionica AM, Matei IA, D'Amico G, Gherman CM, Dom A C, Mihalca AD. Diversity of Flea (Siphonaptera) Parasites on Red Foxes (Vulpes vulpes) in Romania. J Med Entomol. 2017 Sep 1;54(5):1243-1250. doi: 10.1093/jme/tjx067.
Sándor AD, D'Amico G, Gherman CM, Dumitrache MO, Domșa C, Mihalca AD. Mesocarnivores and macroparasites: altitude and land use predict the ticks occurring on red foxes (Vulpes vulpes). Parasit Vectors. 2017 Apr 5;10(1):173. doi: 10.1186/s13071-017-2113-9.
D'Amico G, Dumitrache MO, Matei IA, Ionică AM, Gherman CM, Sándor AD, Modrý D, Mihalca AD. Ixodid ticks parasitizing wild carnivores in Romania. Exp Appl Acarol. 2017 Feb;71(2):139-149. doi: 10.1007/s10493-017-0108-z.
Hornok S, Sándor AD, Beck R, Farkas R, Beati L, Kontschán J, Takács N, Földvári G, Silaghi C, Meyer-Kayser E, Hodžić A, Tomanović S, Abdullah S, Wall R, Estrada-Peña A, Duscher GG, Plantard O. Contributions to the phylogeny of Ixodes (Pholeoixodes) canisuga, I. (Ph.) kaiseri, I. (Ph.) hexagonus and a simple pictorial key for the identification of their females. Parasit Vectors. 2017 Nov 3;10(1):545. doi: 10.1186/s13071-017-2424-x.
Rigó K, Majoros G, Szekeres S, Molnár I, Jablonszky M, Majláthová V, Majláth I, Földvári G. Identification of Hepatozoon erhardovae Krampitz, 1964 from bank voles (Myodes glareolus) and fleas in Southern Hungary. Parasitol Res. 2016 Jun;115(6):2409-13. doi: 10.1007/s00436-016-4992-7.
Hornok S, Trauttwein K, Takács N, Hodžić A, Duscher GG, Kontschán J. Molecular analysis of Ixodes rugicollis, Candidatus Neoehrlichia sp. (FU98) and a novel Babesia genotype from a European badger (Meles meles). Ticks Tick Borne Dis. 2017 Jan;8(1):41-44. doi: 10.1016/j.ttbdis.2016.09.014.
Hornok S, Wang Y, Otranto D, Keskin A, Lia RP, Kontschán J, Takács N, Farkas R, Sándor AD. Phylogenetic analysis of Haemaphysalis erinacei Pavesi, 1884 (Acari: Ixodidae) from China, Turkey, Italy and Romania. Parasit Vectors. 2016 Dec 15;9(1):643. doi: 10.1186/s13071-016-1927-1.
Dumitrache MO, Matei IA, Ionică AM, Kalmár Z, D'Amico G, Sikó-Barabási S, Ionescu DT, Gherman CM, Mihalca AD. Molecular detection of Anaplasma phagocytophilum and Borrelia burgdorferi sensu lato genospecies in red foxes (Vulpes vulpes) from Romania. Parasit Vectors. 2015 Oct 8;8:514. doi: 10.1186/s13071-015-1130-9.
Szekeres S, Claudia Coipan E, Rigó K, Majoros G, Jahfari S, Sprong H, Földvári G. Candidatus Neoehrlichia mikurensis and Anaplasma phagocytophilum in natural rodent and tick communities in Southern Hungary. Ticks Tick Borne Dis. 2015 Mar;6(2):111-6. doi: 10.1016/j.ttbdis.2014.10.004.
Farkas R, Takács N, Hornyák Á, Nachum-Biala Y, Hornok S, Baneth G. First report on Babesia cf. microti infection of red foxes (Vulpes vulpes) from Hungary. Parasit Vectors. 2015 Jan 27;8:55. doi: 10.1186/s13071-015-0660-5.
Farkas R, Solymosi N, Takács N, Hornyák Á, Hornok S, Nachum-Biala Y, Baneth G. First molecular evidence of Hepatozoon canis infection in red foxes and golden jackals from Hungary. Parasit Vectors. 2014 Jul 2;7:303. doi: 10.1186/1756-3305-7-303.
Hornok S, Tánczos B, Fernández de Mera IG, de la Fuente J, Hofmann-Lehmann R, Farkas R. High prevalence of Hepatozoon-infection among shepherd dogs in a region considered to be free of Rhipicephalus sanguineus. Vet Parasitol. 2013 Sep 1;196(1-2):189-93. doi: 10.1016/j.vetpar.2013.02.009.
Gherman CM, Sándor AD, Kalmár Z, Marinov M, Mihalca AD. First report of Borrelia burgdorferi sensu lato in two threatened carnivores: the marbled polecat, Vormela peregusna and the European mink, Mustela lutreola (Mammalia: Mustelidae). BMC Vet Res. 2012 Aug 18;8:137. doi: 10.1186/1746-6148-8-137.
Boretti FS, Perreten A, Meli ML, Cattori V, Willi B, Wengi N, Hornok S, Honegger H, Hegglin D, Woelfel R, Reusch CE, Lutz H, Hofmann-Lehmann R. Molecular Investigations of Rickettsia helvetica infection in dogs, foxes, humans, and Ixodes ticks. Appl Environ Microbiol. 2009 May;75(10):3230-7. doi: 10.1128/AEM.00220-09.
