Research findings

Szekeres, Sándor, Nagy, Gábor, Csivincsik, Ágnes, Takács, Nóra, Kontschán, Jenő, Hornok, Sándor, The European Badger (Meles meles) as a Host for Ticks and Tick-Borne Pathogens in Peri-Urban Environments, Hungary, Transboundary and Emerging Diseases, 2025, 1587951, 10 pages, 2025. https://doi.org/10.1155/tbed/1587951

Sándor AD, Domșa C, Péter Á, Hornok S. Ixodid ticks of Western Palearctic bats: ecology, host-parasite relationships, geographic distribution and zoonotic importance. Front Vet Sci. 2025 Jun 18;12:1517704. doi: 10.3389/fvets.2025.1517704.

Sándor AD, Péter Á, Wong JB, Burri R, Hornok S. Hyalomma rufipes of Asian origin transported to Europe by a migrant bird. Ticks Tick Borne Dis. 2025 Sep;16(5):102518. doi: 10.1016/j.ttbdis.2025.102518.

Péter Á, Beke B, László Z, Hornok S, Sándor AD. Contrasting effects of body condition on ectoparasite abundance in a social bat: different roles of season and host sex. Int J Parasitol. 2025 Aug;55(10):537-546. doi: 10.1016/j.ijpara.2025.04.015.

Ngoc DP, Ha TDT, Ngoc SN, Ngoc MP, Nhu ND, Farkas R, Teeraananchai S, Hornok S. The current status and predicted climate-driven range expansion of Rhipicephalus microplus in northern Vietnam. Acta Trop. 2025 Aug;268:107732. doi: 10.1016/j.actatropica.2025.107732.

Lakos A, Nagy G, Varga Z, Rásonyi-Kovács P, Hornok S. Reassessing Chronic Lyme Disease and Post-Treatment Lyme Disease Syndrome as Focal Infections. Cureus. 2025 Aug 29;17(8):e91246. doi: 10.7759/cureus.91246.

Kürtös G, Rácz N, Kassay V, Olaszy K, Vizi Z, Szilasi A, Takács N, Keve G, Tuska-Szalay B, Hornok S. Infection caused by a parasite most closely related to Hepatozoon luiperdjie in cats from Hungary. Ticks Tick Borne Dis. 2025 Nov;16(6):102556. doi: 10.1016/j.ttbdis.2025.102556.

Keve G, Varga T, Hoitsy M, Wermer K, Hornok S. An assessment of the possible recent establishment of Hyalomma rufipes in Hungary. Ticks Tick Borne Dis. 2025 Sep;16(5):102544. doi: 10.1016/j.ttbdis.2025.102544.

Keve G, Sándor AD, Hornok S. A review on the role of birds as disseminators of ticks, with special emphasis on Hyalomma species and Crimean-Congo haemorrhagic fever virus. Ticks Tick Borne Dis. 2025 Sep;16(5):102535. doi: 10.1016/j.ttbdis.2025.102535. Epub 2025 Aug 28. Erratum in: Ticks Tick Borne Dis. 2026 Jan 16:102587. doi: 10.1016/j.ttbdis.2025.102587.

Keve G, Reynolds C, Takács N, Hornok S. Uneven temporal distribution of piroplasms (Piroplasmida: Babesiidae, Theileriidae) in Haemaphysalis concinna in an urban biotope of the Western Palearctic focus region of this tick species. Ticks Tick Borne Dis. 2025 Mar;16(2):102458. doi: 10.1016/j.ttbdis.2025.102458.

Hornok S, Keskin A, Uspensky I, Kontschán J, Takács N, Lesiczka P, Warbroek T, van den Bosch TJM, Keve G, Pitó A, Sándor AD. Updates on subgenus Ixodes in the Mediterranean region: validity of Ixodes festai Rondelli, 1926, reinstatement of Ixodes tatei Arthur, 1959, and a new species closely related to Ixodes gibbosus Nuttall, 1916. Int J Parasitol. 2025 Sep 19:S0020-7519(25)00177-8. doi: 10.1016/j.ijpara.2025.09.002.

Hornok S, Takács N, Nagy G, Lakos A. Retrospective molecular analyses of hard ticks (Acari: Ixodidae) from patients admitted to the Centre for Tick-Borne Diseases in Central Europe, Hungary (1999-2021), in relation to clinical symptoms. Parasit Vectors. 2025 Jun 20;18(1):229. doi: 10.1186/s13071-025-06880-2.

Hornok S, Keve G, Tuska-Szalay B. Transmission route-dependent genetic diversity of selected protozoan parasites as reflected by the phylogenetic analysis of the 18S rRNA gene. Acta Vet Hung. 2025 Mar 27;73(1):64-72. doi: 10.1556/004.2025.01128.

Hornok S, Kontschán J, Keve G, Takács N, Van Nguyen D, Ho KNP, Görföl T, Wang Y, Farkas R, Dao TTH. First report of Haemaphysalis bispinosa, molecular-geographic relationships of Ixodes granulatus and a new Dermacentor species from Vietnam. Parasit Vectors. 2025 Jan 23;18(1):21. doi: 10.1186/s13071-024-06641-7.

Hornok S, Lesiczka P, Warbroek T, van den Bosch TJM, Pitó A, Keve G, Takács N, Kontschán J, Sándor AD. Comparison and complete mitogenomes of two morphologically similar but ecologically different tick species, Ixodes arboricola and Ixodes lividus (subgenus Pholeoixodes). Ticks Tick Borne Dis. 2026 Jan 10;17(1):102589. doi: 10.1016/j.ttbdis.2025.102589.

Gál J, Ziszisz Á, Hoitsy M, Sós E, Hornok S, Marosán M, Tóth T, Mándoki M, Veprik R, Könyves L, Takács N, Kaszab E, Bali K, Keve G. First detection of Ehrlichia cf. ewingii in a reindeer (Rangifer tarandus) in Hungary. Acta Vet Hung. 2025 Jul 17;73(3):144-149. doi: 10.1556/004.2025.01151. 

Bendjeddou ML, Khelfaoui F, Abiadh A, Mechouk N, Mihalca AD, Sándor AD. Bat Ectoparasites (Acari, Diptera, Hemiptera, Siphonaptera) in the Grand Maghreb (Algeria, Libya, Mauritania, Morocco and Tunisia): A Literature Review and New Data. Acta Parasitol. 2024 Mar;69(1):106-120. doi: 10.1007/s11686-023-00732-8.

Ganzinelli S, Hamšíková Z, Földvári G, Szekeres S, Pfeffer M, Schnittger L, Kazimírová M. Phylogenetic analysis of a novel Hepatozoon species (Hepatozoon sp. SK3) and an additional yet unknown Hepatozoon species (Hepatozoon sp. BV2) besides H. erhardovae in small rodents from Central Europe. Parasitol Res. 2024 Jun 24;123(6):250. doi: 10.1007/s00436-024-08269-z

Wu X, Xu J, Su L, Li E, Wang S, Hornok S, Liu G, Wang Y. First Molecular Evidence of Babesia caballi and Theileria equi in Imported Donkeys from Kyrgyzstan. Pathogens. 2024 Aug 23;13(9):713. doi: 10.3390/pathogens13090713.

Szentiványi T, Takács N, Sándor AD, Péter Á, Boldogh SA, Kováts D, Foster JT, Estók P, Hornok S. Bat-associated ticks as a potential link for vector-borne pathogen transmission between bats and other animals. PLoS Negl Trop Dis. 2024 Oct 25;18(10):e0012584. doi: 10.1371/journal.pntd.0012584.

Pitó A, Fedorov D, Brlík V, Kontschán J, Keve G, Sándor AD, Takács N, Hornok S. East-to-west dispersal of bird-associated ixodid ticks in the northern Palaearctic: Review of already reported tick species according to longitudinal migratory avian hosts and first evidence on the genetic connectedness of Ixodes apronophorus between Siberia and Europe. Curr Res Parasitol Vector Borne Dis. 2024 Jul 14;6:100201. doi: 10.1016/j.crpvbd.2024.100201.

Li E, Wu X, Tang L, Yang M, Hornok S, Zhang C, Zhang Y, Zhao G, Wang Y. Molecular-phylogenetic analyses of Babesia and Theileria species from small mammals and their ticks in northern China suggest new reservoirs of bovine and equine piroplasms. Vet Parasitol. 2024 Dec;332:110304. doi: 10.1016/j.vetpar.2024.110304.

Hornok S, Kontschán J, Takács N, Pap PL, Sándor AD. First record of Ixodes (Scaphixodes) caledonicus in the Carpathian Basin and first time molecular-phylogenetic analysis of this tick species with updated host records and geographical range. Ticks Tick Borne Dis. 2024 Jan;15(1):102280. doi: 10.1016/j.ttbdis.2023.102280.

Hornok S, Farkas R, Duong NN, Kontschán J, Takács N, Keve G, Pham DN, Dao TTH. A morpho phylogenetic update on ixodid ticks infesting cattle and buffalos in Vietnam, with three new species to the fauna and a checklist of all species indigenous to the country. Parasit Vectors. 2024 Jul 25;17(1):319. doi: 10.1186/s13071-024-06384-5

Hornok S, Kontschán J, Takano A, Gotoh Y, Hassanin A, Tu VT. Description of Ixodes lanigeri sp. nov., a new hard tick species (Acari, Ixodidae) collected from mouse-eared bats (Vespertilionidae, Myotis) in Vietnam. Zookeys. 2024 Oct 14;1215:107-125. doi: 10.3897/zookeys.1215.123624.

Dao TTH, Takács N, Tran TN, Truong AN, Skinner K, Kontschán J, Farkas R, Hornok S. Detection of tick- borne pathogens in the pangolin tick, Amblyomma javanense, from Vietnam and Laos, including a novel species of Trypanosoma. Acta Trop. 2024 Dec;260:107384. doi: 10.1016/j.actatropica.2024.107384. Epub 2024 Sep 10.

Cui N, Su L, Wang Z, Hornok S, Tang L, Yang M, Zhang Y, Zhao G, Wang Y. First identification of Cytauxzoon manul in Eurasian lynx (Lynx lynx) in northwestern China. Parasit Vectors. 2024 Jun 6;17(1):249. doi: 10.1186/s13071-024-06326-1.

Pitó A, Bukor B, Győrig E, Brlík V, Kontschán J, Keve G, Takács N, Hornok, S. (2024) Investigations of the tick burden on passeriform, water-associated and predatory birds reveal new tick–host associations and habitat-related factors of tick infestation. Parasites and Vectors 17: 1 Paper: 144. https://parasitesandvectors.biomedcentral.com/articles/10.1186/s13071-024-06229-1

Liu G, Tan W, Wang H, Han X, Hornok S, Zhao S, Mi L, Wang S, Yang M, Wang Y. (2024) The great gerbil (Rhombomys opimus) as a host for tick species in Gurbantunggut Desert. Parasites and Vectors 17: 1 Paper: 55, 7 p.
https://parasitesandvectors.biomedcentral.com/articles/10.1186/s13071-024-06160-5

Keve G, Csörgő T, Kováts D, Hornok S. (2024) Long term evaluation of factors influencing the association of ixodid ticks with birds in Central Europe, Hungary. Scientific Reports 14: 1 Paper: 4958.
https://www.nature.com/articles/s41598-024-55021-9

Halajian A, Cuozzo FP, Heyne H, Sauther ML, Linden B, Linden J, Tordiffe ASW, Rampedi KM, Hornok S. (2024) Hard ticks (Acari: Ixodidae) parasitizing bushbabies (Mammalia: Galagidae) in a biodiversity hotspot of northern South Africa. Ticks and Tick-Borne Diseases 15:2.102313 p.
https://doi.org/10.1016/j.ttbdis.2024.102313

Fedorov D, Hornok S. (2024) Checklist of hosts, illustrated geographical range, and ecology of tick species from the genus Ixodes (Acari, Ixodidae) in Russia and other post-Soviet countries.
Zookeys 1201: pp. 255-343., 89 p. https://zookeys.pensoft.net/article/115467/

Bendjeddou ML, Khelfaoui F, Abiadh A, Sándor AD et al (2023) Bat Ectoparasites (Acari, Diptera, Hemiptera, Siphonaptera) in the Grand Maghreb (Algeria, Libya, Mauritania, Morocco and Tunisia): A Literature Review and New Data. Acta Parasitologica 11. https://doi.org/10.1007/s11686-023-00732-8

Matei IA, Kalmár Z, Balea A, Mihaiu M, Sándor AD, Cocian A, Crăciun S, Bouari C, Briciu VT, Fiț N (2023) The role of wild boars in the circulation of some tick-borne pathogens: the first evidence of Rickettsia monacensis presence. Animals 13: 1743. doi: 10.3390/ani13111743, IF: 3.0

Tuska-Szalay B, Boldogh SA, Farkas R, Rompos L, Takács N, Beresnyák V, Hornok S (2023). Screening of Domestic Cats from North-Eastern Hungary for Hepatozoon felis and Cytauxzoon europaeus That Cause Infections in Local Wildcat Populations. Pathogens 12(5): 656. http://doi.org/10.3390/pathogens12050656

Takano A, Yamauchi T, Takahashi M, Shimoda H, Gotoh Y, Mizuno J, Hornok S. (2023). Description of three new bat-associated species of hard ticks (Acari, Ixodidae) from Japan. Zookeys 1180: 1–26. http://doi.org/10.3897/zookeys.1180.108418

Hornok S, Kontschán J, Takács N, Heyne H, Kovács ÁB, Plantard O, Halajian A et al. (2023). Molecular-phylogenetic analyses of Ixodes species from South Africa suggest an African origin of bird-associated exophilic ticks (subgenus Trichotoixodes). Parasites and Vectors 16(1). http://doi.org/10.1186/s13071-023-05998-5

Hornok S, Kontschán J, Takács N, Pap PL, Sándor AD (2024). First record of Ixodes (Scaphixodes) caledonicus in the Carpathian Basin and first time molecular-phylogenetic analysis of this tick species with updated host records and geographical range. Ticks and Tick-borne Diseases 15(1): 102280. http://doi.org/10.1016/j.ttbdis.2023.102280

Matei I, Kalmár Zs, Balea A, Mihaiu M, Sándor AD, Cocian A, Craciun S, Bouari C, Briciu V, Fit N. The Role of Wild Boars in the Circulation of Tick-Borne Pathogens: The First Evidence of Rickettsia monacensis Presence. Animals. 2023 May;13 (11), 1743. doi: 10.3390/ani13111743.

Hornok S, Mihalca AD, Kontschán J, Takács N, Fedorov D, Plantard O, Sándor AD. Phylogenetic analyses of Ixodes rugicollis with notes on its morphology in comparison with Ixodes cornutus. Parasit Vectors. 2023 Mar 16;16(1):106. doi: 10.1186/s13071-023-05718-z.

Keve G, Csörgő T, Benke A, Huber A, Mórocz A, Németh Á, Kalocsa B, Tamás EA, Gyurácz J, Kiss O, Kováts D, Sándor AD, Karcza Z, Hornok S. Ornithological and molecular evidence of a reproducing Hyalomma rufipes population under continental climate in Europe. Front Vet Sci. 2023 Mar 22;10:1147186. doi: 10.3389/fvets.2023.1147186.

Ivan T, Matei IA, Novac CȘ, Kalmár Z, Borșan SD, Panait LC, Gherman CM, Ionică AM, Papuc I, Mihalca AD. Spotted Fever Group Rickettsia spp. Diversity in Ticks and the First Report of Rickettsia hoogstraalii in Romania. Vet Sci. 2022 Jul 8;9(7):343. doi: 10.3390/vetsci9070343.

Hornok S, Szekeres S, Horváth G, Takács N, Bekő K, Kontschán J, Gyuranecz M, Tóth B, Sándor AD, Juhász A, Beck R, Farkas R. Diversity of tick species and associated pathogens on peri-urban wild boars - First report of the zoonotic Babesia cf. crassa from Hungary. Ticks Tick Borne Dis. 2022 May;13(3):101936. doi: 10.1016/j.ttbdis.2022.101936.

Hornok S, Boldogh SA, Takács N, Kontschán J, Szekeres S, Sós E, Sándor AD, Wang Y, Tuska-Szalay B. Molecular epidemiological study on ticks and tick-borne protozoan parasites (Apicomplexa: Cytauxzoon and Hepatozoon spp.) from wild cats (Felis silvestris), Mustelidae and red squirrels (Sciurus vulgaris) in central Europe, Hungary. Parasit Vectors. 2022 May 21;15(1):174. doi: 10.1186/s13071-022-05271-1.

Hornok S, Cutajar B, Takács N, Galea N, Attard D, Coleiro C, Galea R, Keve G, Sándor AD, Kontschán J. On the way between Africa and Europe: Molecular taxonomy of ticks collected from birds in Malta. Ticks Tick Borne Dis. 2022 Sep;13(5):102001. doi: 10.1016/j.ttbdis.2022.102001.

Hornok S, Boldogh SA, Takács N, Sándor AD, Tuska-Szalay B. Zoonotic ecotype-I of Anaplasma phagocytophilum in sympatric wildcat, pine marten and red squirrel - Short communication. Acta Vet Hung. 2022 Aug 26. doi: 10.1556/004.2022.00021.

Keve G, Sándor AD, Hornok S. Hard ticks (Acari: Ixodidae) associated with birds in Europe: Review of literature data. Front Vet Sci. 2022 Aug 25;9:928756. doi: 10.3389/fvets.2022.928756.

Matei IA, Corduneanu A, Sándor AD, Ionică AM, Panait L, Kalmár Z, Ivan T, Papuc I, Bouari C, Fit N, Mihalca AD. Rickettsia spp. in bats of Romania: high prevalence of Rickettsia monacensis in two insectivorous bat species. Parasit Vectors. 2021 Feb 10;14(1):107. doi: 10.1186/s13071-021-04592-x.

Matei IA, Ionică AM, Corduneanu A, Domșa C, Sándor AD. The Presence of Ehrlichia Canis in Rhipicephalus Bursa Ticks Collected from Ungulates in Continental Eastern Europe. J Vet Res. 2021 Sep 23;65(3):271-275. doi: 10.2478/jvetres-2021-0044.

Sándor AD, Péter Á, Corduneanu A, Barti L, Csősz I, Kalmár Z, Hornok S, Kontschán J, Mihalca AD. Wide Distribution and Diversity of Malaria-Related Haemosporidian Parasites (Polychromophilus spp.) in Bats and Their Ectoparasites in Eastern Europe. Microorganisms. 2021 Jan 22;9(2):230. doi: 10.3390/microorganisms9020230.

Hornok S, Meyer-Kayser E, Kontschán J, Takács N, Plantard O, Cullen S, Gaughran A, Szekeres S, Majoros G, Beck R, Boldogh SA, Horváth G, Kutasi C, Sándor AD. Morphology of Pholeoixodes species associated with carnivores in the western Palearctic: Pictorial key based on molecularly identified Ixodes (Ph.) canisuga, I. (Ph.) hexagonus and I. (Ph.) kaiseri males, nymphs and larvae. Ticks Tick Borne Dis. 2021 Jul;12(4):101715. doi: 10.1016/j.ttbdis.2021.101715.

Sang C, Yang Y, Dong Q, Xu B, Liu G, Hornok S, Liu Z, Wang Y, Hazihan W. Molecular survey of Babesia spp. in red foxes (Vulpes Vulpes), Asian badgers (Meles leucurus) and their ticks in China. Ticks Tick Borne Dis. 2021 Jul;12(4):101710. doi: 10.1016/j.ttbdis.2021.101710.

Péter Á, Barti L, Corduneanu A, Hornok S, Mihalca AD, Sándor AD. First record of Ixodes simplex found on a human host, with a review of cases of human infestation by bat tick species occurring in Europe. Ticks Tick Borne Dis. 2021 Jul;12(4):101722. doi: 10.1016/j.ttbdis.2021.101722.

Liu G, Zhao S, Tan W, Hornok S, Yuan W, Mi L, Wang S, Liu Z, Zhang Y, Hazihan W, Gu X, Wang Y. Rickettsiae in red fox (Vulpes vulpes), marbled polecat (Vormela peregusna) and their ticks in northwestern China. Parasit Vectors. 2021 Apr 19;14(1):204. doi: 10.1186/s13071-021-04718-1.

Sándor AD, Milchev B, Takács N, Kontschán J, Szekeres S, Hornok S. Five ixodid tick species including two morphotypes of Rhipicephalus turanicus on nestlings of Eurasian eagle owl (Bubo bubo) from south-eastern Bulgaria. Parasit Vectors. 2021 Jun 26;14(1):334. doi: 10.1186/s13071-021-04832-0.

Hornok S, Kontschán J, Takács N, Chaber AL, Halajian A, Szekeres S, Sándor AD, Plantard O. Rickettsiaceae in two reptile-associated tick species, Amblyomma exornatum and Africaniella transversale: First evidence of Occidentia massiliensis in hard ticks (Acari: Ixodidae). Ticks Tick Borne Dis. 2022 Jan;13(1):101830. doi: 10.1016/j.ttbdis.2021.101830.

Hornok S, Daccord J, Takács N, Kontschán J, Tuska-Szalay B, Sándor AD, Szekeres S, Meli ML, Hofmann-Lehmann R. Investigation on haplotypes of ixodid ticks and retrospective finding of Borrelia miyamotoi in bank vole (Myodes glareolus) in Switzerland. Ticks Tick Borne Dis. 2022 Jan;13(1):101865. doi: 10.1016/j.ttbdis.2021.101865.

Tuska-Szalay B, Vizi Z, Hofmann-Lehmann R, Vajdovich P, Takács N, Meli ML, Farkas R, Stummer-Knyihár V, Jerzsele Á, Kontschán J, Szekeres S, Hornok S. Babesia gibsoni emerging with high prevalence and co-infections in "fighting dogs" in Hungary. Curr Res Parasitol Vector Borne Dis. 2021 Sep 24;1:100048. doi: 10.1016/j.crpvbd.2021.100048.

Kalmár Z, Dumitrache MO, D'Amico G, Matei IA, Ionică AM, Gherman CM, Lupșe M, Mihalca AD. Multiple Tick-Borne Pathogens in Ixodes ricinus Ticks Collected from Humans in Romania. Pathogens. 2020 May 19;9(5):390. doi: 10.3390/pathogens9050390.

Borşan SD, Toma-Naic A, Péter Á, Sándor AD, Peștean C, Mihalca AD. Impact of abiotic factors, habitat type and urban wildlife on the ecology of hard ticks (Acari: Ixodidae) in urban and peri-urban habitats. Parasit Vectors. 2020 Sep 18;13(1):476. doi: 10.1186/s13071-020-04352-3.

Hornok S, Grima A, Takács N, Szekeres S, Kontschán J. First records and molecular-phylogenetic analyses of three tick species (Ixodes kaiseri, Hyalomma lusitanicum and Ornithodoros coniceps) from Malta. Ticks Tick Borne Dis. 2020 May;11(3):101379. doi: 10.1016/j.ttbdis.2020.101379.

Liu X, Yang M, Song S, Liu G, Zhao S, Liu G, Hornok S, Wang Y, Jiang H. Brucella melitensis in Asian Badgers, Northwestern China. Emerg Infect Dis. 2020 Apr;26(4):804-806. doi: 10.3201/eid2604.190833.

Yan B, Zhu Q, Xu J, Zhao S, Piao D, Chen C, Liu Q, Hornok S, Wang Y, Wang B, Jiang H. Brucella in Himalayan Marmots (Marmota himalayana). J Wildl Dis. 2020 Jul;56(3):730-732. doi: 10.7589/2019-09-237.

Hornok S, Boldogh SA, Takács N, Juhász A, Kontschán J, Földi D, Koleszár B, Morandini P, Gyuranecz M, Szekeres S. Anaplasmataceae closely related to Ehrlichia chaffeensis and Neorickettsia helminthoeca from birds in Central Europe, Hungary. Antonie Van Leeuwenhoek. 2020 Jul;113(7):1067-1073. doi: 10.1007/s10482-020-01415-4.

Hornok S, Sándor AD, Földvári G, Ionică AM, Silaghi C, Takács N, Schötta AM, Wijnveld M. First broad-range molecular screening of tick-borne pathogens in Ixodes (Pholeoixodes) kaiseri, with special emphasis on piroplasms. Acta Vet Hung. 2020 May 8;68(1):30-33. doi: 10.1556/004.2020.00003.

Hornok S, Mühldorfer K, Takács N, Hofmann-Lehmann R, Meli ML, Gyuranecz M, Unnsteinsdóttir ER, Greenwood AD, Czirják GÁ. Broad Range Screening of Vector-Borne Pathogens in Arctic Foxes (Vulpes lagopus) in Iceland. Animals (Basel). 2020 Nov 4;10(11):2031. doi: 10.3390/ani10112031.

McKee CD, Krawczyk AI, Sándor AD, Görföl T, Földvári M, Földvári G, Dekeukeleire D, Haarsma A-, Kosoy MY, Webb CT, Sprong H. Host phylogeny, geographic overlap, and roost sharing shape parasite communities in european bats. Front Ecol Evol 2019 Marc;7(69). doi: 10.3389/fevo.2019.00069

Kalmár Z, Sándor AD, Matei IA, Ionică A, D'Amico G, Gherman CM, Mihalca AD. Borrelia spp. in small mammals in Romania. Parasit Vectors. 2019 Sep 24;12(1):461. doi: 10.1186/s13071-019-3713-3.

Hornok S, Szőke K, Meli ML, Sándor AD, Görföl T, Estók P, Wang Y, Tu VT, Kováts D, Boldogh SA, Corduneanu A, Sulyok KM, Gyuranecz M, Kontschán J, Takács N, Halajian A, Epis S, Hofmann-Lehmann R. Molecular detection of vector-borne bacteria in bat ticks (Acari: Ixodidae, Argasidae) from eight countries of the Old and New Worlds. Parasit Vectors. 2019 Jan 22;12(1):50. doi: 10.1186/s13071-019-3303-4.

Zhao S, Yang M, Jiang M, Yan B, Zhao S, Yuan W, Wang B, Hornok S, Wang Y. Rickettsia raoultii and Rickettsia sibirica in ticks from the long-tailed ground squirrel near the China-Kazakhstan border. Exp Appl Acarol. 2019 Mar;77(3):425-433. doi: 10.1007/s10493-019-00349-5.

Zeb J, Szekeres S, Takács N, Kontschán J, Shams S, Ayaz S, Hornok S. Genetic diversity, piroplasms and trypanosomes in Rhipicephalus microplus and Hyalomma anatolicum collected from cattle in northern Pakistan. Exp Appl Acarol. 2019 Oct;79(2):233-243. doi: 10.1007/s10493-019-00418-9.

Norte AC, Margos G, Becker NS, Albino Ramos J, Núncio MS, Fingerle V, Araújo PM, Adamík P, Alivizatos H, Barba E, Barrientos R, Cauchard L, Csörgő T, Diakou A, Dingemanse NJ, Doligez B, Dubiec A, Eeva T, Flaisz B, Grim T, Hau M, Heylen D, Hornok S, Kazantzidis S, Kováts D, Krause F, Literak I, Mänd R, Mentesana L, Morinay J, Mutanen M, Neto JM, Nováková M, Sanz JJ, Pascoal da Silva L, Sprong H, Tirri IS, Török J, Trilar T, Tyller Z, Visser ME, Lopes de Carvalho I. Host dispersal shapes the population structure of a tick-borne bacterial pathogen. Mol Ecol. 2020 Feb;29(3):485-501. doi: 10.1111/mec.15336.

Sándor AD, Corduneanu A, Péter Á, Mihalca AD, Barti L, Csősz I, Szőke K, Hornok S. Bats and ticks: host selection and seasonality of bat-specialist ticks in eastern Europe. Parasit Vectors. 2019 Dec 27;12(1):605. doi: 10.1186/s13071-019-3861-5.

Matei IA, D'Amico G, Ionică AM, Kalmár Z, Corduneanu A, Sándor AD, Fiţ N, Bogdan L, Gherman CM, Mihalca AD. New records for Anaplasma phagocytophilum infection in small mammal species. Parasit Vectors. 2018 Mar 20;11(1):193. doi: 10.1186/s13071-018-2791-y.

Hornok S, Szőke K, Estók P, Krawczyk A, Haarsma AJ, Kováts D, Boldogh SA, Morandini P, Szekeres S, Takács N, Kontschán J, Meli ML, Fernández de Mera IG, de la Fuente J, Gyuranecz M, Sulyok KM, Weibel B, Gönczi E, de Bruin A, Sprong H, Hofmann-Lehmann R. Assessing bat droppings and predatory bird pellets for vector-borne bacteria: molecular evidence of bat-associated Neorickettsia sp. in Europe. Antonie Van Leeuwenhoek. 2018 Sep;111(9):1707-1717. doi: 10.1007/s10482-018-1043-7.

Hornok S, Sugár L, Fernández de Mera IG, de la Fuente J, Horváth G, Kovács T, Micsutka A, Gönczi E, Flaisz B, Takács N, Farkas R, Meli ML, Hofmann-Lehmann R. Tick- and fly-borne bacteria in ungulates: the prevalence of Anaplasma phagocytophilum, haemoplasmas and rickettsiae in water buffalo and deer species in Central Europe, Hungary. BMC Vet Res. 2018 Mar 20;14(1):98. doi: 10.1186/s12917-018-1403-6.

Hornok S, Corduneanu A, Kontschán J, Bekő K, Szőke K, Görföl T, Gyuranecz M, Sándor AD. Analyses of separate and concatenated cox1 and 18S rRNA gene sequences indicate that the bat piroplasm Babesia vesperuginis is phylogenetically close to Cytauxzoon felis and the 'prototheilerid' Babesia conradae. Acta Vet Hung. 2018 Mar;66(1):107-115. doi: 10.1556/004.2018.010.

Hornok S, Horváth G, Takács N, Kontschán J, Szőke K, Farkas R. Molecular identification of badger-associated Babesia sp. DNA in dogs: updated phylogeny of piroplasms infecting Caniformia. Parasit Vectors. 2018 Apr 11;11(1):235. doi: 10.1186/s13071-018-2794-8.

Hornok S, Grima A, Takács N, Kontschán J. Infestation of Rhipicephalus sanguineus sensu lato on cats in Malta. Ticks Tick Borne Dis. 2018 Jul;9(5):1120-1124. doi: 10.1016/j.ttbdis.2018.04.007.

Hornok S, Horváth G, Takács N, Farkas R, Szőke K, Kontschán J. Molecular evidence of a badger-associated Ehrlichia sp., a Candidatus Neoehrlichia lotoris-like genotype and Anaplasma marginale in dogs. Ticks Tick Borne Dis. 2018 Jul;9(5):1302-1309. doi: 10.1016/j.ttbdis.2018.05.012.

Sándor AD, Kontschán J, Plantard O, Péter Á, Hornok S. Illustrated redescription of the male of Ixodes simplex Neumann, 1906. Ticks Tick Borne Dis. 2018 Jul;9(5):1328-1330. doi: 10.1016/j.ttbdis.2018.05.011.

Liu X, Yang M, Liu G, Zhao S, Yuan W, Xiao R, Hazihan W, Hornok S, Wang Y. Molecular evidence of Rickettsia raoultii, "Candidatus Rickettsia barbariae" and a novel Babesia genotype in marbled polecats (Vormela peregusna) at the China-Kazakhstan border. Parasit Vectors. 2018 Aug 4;11(1):450. doi: 10.1186/s13071-018-3033-z.

Matei IA, Kalmár Z, Lupşe M, D'Amico G, Ionică AM, Dumitrache MO, Gherman CM, Mihalca AD. The risk of exposure to rickettsial infections and human granulocytic anaplasmosis associated with Ixodes ricinus tick bites in humans in Romania: A multiannual study. Ticks Tick Borne Dis. 2017 Mar;8(3):375-378. doi: 10.1016/j.ttbdis.2016.12.013.

Sándor AD, D'Amico G, Gherman CM, Dumitrache MO, Domșa C, Mihalca AD. Mesocarnivores and macroparasites: altitude and land use predict the ticks occurring on red foxes (Vulpes vulpes). Parasit Vectors. 2017 Apr 5;10(1):173. doi: 10.1186/s13071-017-2113-9.

D'Amico G, Dumitrache MO, Matei IA, Ionică AM, Gherman CM, Sándor AD, Modrý D, Mihalca AD. Ixodid ticks parasitizing wild carnivores in Romania. Exp Appl Acarol. 2017 Feb;71(2):139-149. doi: 10.1007/s10493-017-0108-z.

Flaisz B, Sulyok KM, Kováts D, Kontschán J, Csörgő T, Csipak Á, Gyuranecz M, Hornok S. Babesia genotypes in Haemaphysalis concinna collected from birds in Hungary reflect phylogeographic connections with Siberia and the Far East. Ticks Tick Borne Dis. 2017 Jun;8(4):666-670. doi: 10.1016/j.ttbdis.2017.04.013.

Hornok S, Sugár L, Horváth G, Kovács T, Micsutka A, Gönczi E, Flaisz B, Takács N, Farkas R, Meli ML, Hofmann-Lehmann R. Evidence for host specificity of Theileria capreoli genotypes in cervids. Parasit Vectors. 2017 Oct 10;10(1):473. doi: 10.1186/s13071-017-2403-2.

Hornok S, Sándor AD, Beck R, Farkas R, Beati L, Kontschán J, Takács N, Földvári G, Silaghi C, Meyer-Kayser E, Hodžić A, Tomanović S, Abdullah S, Wall R, Estrada-Peña A, Duscher GG, Plantard O. Contributions to the phylogeny of Ixodes (Pholeoixodes) canisuga, I. (Ph.) kaiseri, I. (Ph.) hexagonus and a simple pictorial key for the identification of their females. Parasit Vectors. 2017 Nov 3;10(1):545. doi: 10.1186/s13071-017-2424-x.

Corduneanu A, Hrazdilová K, Sándor AD, Matei IA, Ionică AM, Barti L, Ciocănău MA, Măntoiu DȘ, Coroiu I, Hornok S, Fuehrer HP, Leitner N, Bagó Z, Stefke K, Modrý D, Mihalca AD. Babesia vesperuginis, a neglected piroplasmid: new host and geographical records, and phylogenetic relations. Parasit Vectors. 2017 Dec 6;10(1):598. doi: 10.1186/s13071-017-2536-3.

Szőke K, Sándor AD, Boldogh SA, Görföl T, Votýpka J, Takács N, Estók P, Kováts D, Corduneanu A, Molnár V, Kontschán J, Hornok S. DNA of free-living bodonids (Euglenozoa: Kinetoplastea) in bat ectoparasites: potential relevance to the evolution of parasitic trypanosomatids. Acta Vet Hung. 2017 Dec;65(4):531-540. doi: 10.1556/004.2017.051.

Matei IA, D'Amico G, Yao PK, Ionică AM, Kanyari PW, Daskalaki AA, Dumitrache MO, Sándor AD, Gherman CM, Qablan M, Modrý D, Mihalca AD. Molecular detection of Anaplasma platys infection in free-roaming dogs and ticks from Kenya and Ivory Coast. Parasit Vectors. 2016 Mar 16;9:157. doi: 10.1186/s13071-016-1443-3.

Sándor AD, Kalmár Z, Matei I, Ionică AM, Mărcuţan ID. Urban Breeding Corvids as Disseminators of Ticks and Emerging Tick-Borne Pathogens. Vector Borne Zoonotic Dis. 2017 Feb;17(2):152-154. doi: 10.1089/vbz.2016.2054.

Mărcuţan ID, Kalmár Z, Ionică AM, D'Amico G, Mihalca AD, Vasile C, Sándor AD. Spotted fever group rickettsiae in ticks of migratory birds in Romania. Parasit Vectors. 2016 May 20;9(1):294. doi: 10.1186/s13071-016-1565-7.

Rigó K, Majoros G, Szekeres S, Molnár I, Jablonszky M, Majláthová V, Majláth I, Földvári G. Identification of Hepatozoon erhardovae Krampitz, 1964 from bank voles (Myodes glareolus) and fleas in Southern Hungary. Parasitol Res. 2016 Jun;115(6):2409-13. doi: 10.1007/s00436-016-4992-7.

Hornok S, Flaisz B, Takács N, Kontschán J, Csörgő T, Csipak Á, Jaksa BR, Kováts D. Bird ticks in Hungary reflect western, southern, eastern flyway connections and two genetic lineages of Ixodes frontalis and Haemaphysalis concinna. Parasit Vectors. 2016 Feb 24;9:101. doi: 10.1186/s13071-016-1365-0.

Hornok S, Kováts D, Flaisz B, Csörgő T, Könczöl Á, Balogh GT, Csorba A, Hunyadi A. An unexpected advantage of insectivorism: insect moulting hormones ingested by song birds affect their ticks. Sci Rep. 2016 Mar 21;6:23390. doi: 10.1038/srep23390.

Hornok S, Görföl T, Estók P, Tu VT, Kontschán J. Description of a new tick species, Ixodes collaris n. sp. (Acari: Ixodidae), from bats (Chiroptera: Hipposideridae, Rhinolophidae) in Vietnam. Parasit Vectors. 2016 Jun 10;9(1):332. doi: 10.1186/s13071-016-1608-0.

Hornok S, Kováts D, Angyal D, Dányi L, Kovács R, Kontschán J. Description of the male and the larva of Ixodes ariadnae Hornok, 2014. Ticks Tick Borne Dis. 2016 Oct;7(6):1252-1255. doi: 10.1016/j.ttbdis.2016.07.005.

Hornok S, Trauttwein K, Takács N, Hodžić A, Duscher GG, Kontschán J. Molecular analysis of Ixodes rugicollis, Candidatus Neoehrlichia sp. (FU98) and a novel Babesia genotype from a European badger (Meles meles). Ticks Tick Borne Dis. 2017 Jan;8(1):41-44. doi: 10.1016/j.ttbdis.2016.09.014.

Hornok S, Szőke K, Kováts D, Estók P, Görföl T, Boldogh SA, Takács N, Kontschán J, Földvári G, Barti L, Corduneanu A, Sándor AD. DNA of Piroplasms of Ruminants and Dogs in Ixodid Bat Ticks. PLoS One. 2016 Dec 8;11(12):e0167735. doi: 10.1371/journal.pone.0167735.

Hornok S, Wang Y, Otranto D, Keskin A, Lia RP, Kontschán J, Takács N, Farkas R, Sándor AD. Phylogenetic analysis of Haemaphysalis erinacei Pavesi, 1884 (Acari: Ixodidae) from China, Turkey, Italy and Romania. Parasit Vectors. 2016 Dec 15;9(1):643. doi: 10.1186/s13071-016-1927-1.

Hornok S, Krawczyk A. First record of Ixodes ariadnae in Western Europe, Belgium--Short communication. Acta Vet Hung. 2016 Dec;64(4):467-471. doi: 10.1556/004.2016.043.

Kalmár Z, Cozma V, Sprong H, Jahfari S, D'Amico G, Mărcuțan DI, Ionică AM, Magdaş C, Modrý D, Mihalca AD. Transstadial transmission of Borrelia turcica in Hyalomma aegyptium ticks. PLoS One. 2015 Feb 19;10(2):e0115520. doi: 10.1371/journal.pone.0115520.

Mărcuţan ID, Sándor AD, Kalmár Zs, Mihalca AD, Gherman CM, Cozma V, D'Amico G. Anaplasma phagocytophilum in ticks and tissues collected from wild birds in Romania. Scien Parasit. 2015 Sep 16;3: 103.111. http://scientia.zooparaz.net/2015_16_03/103-111-SP-2015-Marcutan-ORA.pdf

D'Amico G, Dumitrache MO, Široký P, Albrechtová K, Sloboda M, Domşa C, Sándor AD, Balázsi R, Kanyari PW, Modrý D, Mihalca AD. Altitudinal and seasonal differences of tick communities in dogs from pastoralist tribes of Northern Kenya. Vet Parasitol. 2015 Sep 15;212(3-4):318-23. doi: 10.1016/j.vetpar.2015.08.025.

Szekeres S, Claudia Coipan E, Rigó K, Majoros G, Jahfari S, Sprong H, Földvári G. Candidatus Neoehrlichia mikurensis and Anaplasma phagocytophilum in natural rodent and tick communities in Southern Hungary. Ticks Tick Borne Dis. 2015 Mar;6(2):111-6. doi: 10.1016/j.ttbdis.2014.10.004.

Hornok S, Földvári G, Rigó K, Meli ML, Gönczi E, Répási A, Farkas R, Papp I, Kontschán J, Hofmann-Lehmann R. Synanthropic rodents and their ectoparasites as carriers of a novel haemoplasma and vector-borne, zoonotic pathogens indoors. Parasit Vectors. 2015 Jan 15;8:27. doi: 10.1186/s13071-014-0630-3.

Hornok S, Kontschán J, Estrada-Peña A, de Mera IG, Tomanović S, de la Fuente J. Contributions to the morphology and phylogeny of the newly discovered bat tick species, Ixodes ariadnae in comparison with I. vespertilionis and I. simplex. Parasit Vectors. 2015 Jan 24;8:47. doi: 10.1186/s13071-015-0665-0.

Farkas R, Takács N, Hornyák Á, Nachum-Biala Y, Hornok S, Baneth G. First report on Babesia cf. microti infection of red foxes (Vulpes vulpes) from Hungary. Parasit Vectors. 2015 Jan 27;8:55. doi: 10.1186/s13071-015-0660-5.

Hornok S, Estók P, Kováts D, Flaisz B, Takács N, Szőke K, Krawczyk A, Kontschán J, Gyuranecz M, Fedák A, Farkas R, Haarsma AJ, Sprong H. Screening of bat faeces for arthropod-borne apicomplexan protozoa: Babesia canis and Besnoitia besnoiti-like sequences from Chiroptera. Parasit Vectors. 2015 Aug 28;8:441. doi: 10.1186/s13071-015-1052-6.

Hornok S, Estrada-Peña A, Kontschán J, Plantard O, Kunz B, Mihalca AD, Thabah A, Tomanović S, Burazerović J, Takács N, Görföl T, Estók P, Tu VT, Szőke K, Fernández de Mera IG, de la Fuente J, Takahashi M, Yamauchi T, Takano A. High degree of mitochondrial gene heterogeneity in the bat tick species Ixodes vespertilionis, I. ariadnae and I. simplex from Eurasia. Parasit Vectors. 2015 Sep 17;8:457. doi: 10.1186/s13071-015-1056-2.

Hornok S, Takács N, Szőke K, Kunz B. First record of Ixodes ariadnae in Germany - Short communication. Acta Vet Hung. 2015 Sep;63(3):347-51. doi: 10.1556/004.2015.032.

Hornok S, Takács N, Kontschán J, György Z, Micsutka A, Iceton S, Flaisz B, Farkas R, Hofmann-Lehmann R. Diversity of Haemaphysalis-associated piroplasms of ruminants in Central-Eastern Europe, Hungary. Parasit Vectors. 2015 Dec 9;8:627. doi: 10.1186/s13071-015-1236-0.

Sándor AD, Dumitrache MO, D'Amico G, Kiss BJ, Mihalca AD. Rhipicephalus rossicus and not R. sanguineus is the dominant tick species of dogs in the wetlands of the Danube Delta, Romania. Vet Parasitol. 2014 Aug 29;204(3-4):430-2. doi: 10.1016/j.vetpar.2014.05.005.

Dumitrache MO, Kiss B, Dantas-Torres F, Latrofa MS, D'Amico G, Sándor AD, Mihalca AD. Seasonal dynamics of Rhipicephalus rossicus attacking domestic dogs from the steppic region of southeastern Romania. Parasit Vectors. 2014 Mar 10;7:97. doi: 10.1186/1756-3305-7-97.

Sándor AD, Mărcuţan DI, D'Amico G, Gherman CM, Dumitrache MO, Mihalca AD. Do the ticks of birds at an important migratory hotspot reflect the seasonal dynamics of Ixodes ricinus at the migration initiation site? A case study in the Danube Delta. PLoS One. 2014 Feb 19;9(2):e89378. doi: 10.1371/journal.pone.0089378.

Hornok S, Kováts D, Csörgő T, Meli ML, Gönczi E, Hadnagy Z, Takács N, Farkas R, Hofmann-Lehmann R. Birds as potential reservoirs of tick-borne pathogens: first evidence of bacteraemia with Rickettsia helvetica. Parasit Vectors. 2014 Mar 28;7:128. doi: 10.1186/1756-3305-7-128.

Hornok S, Kontschán J, Kováts D, Kovács R, Angyal D, Görföl T, Polacsek Z, Kalmár Z, Mihalca AD. Bat ticks revisited: Ixodes ariadnae sp. nov. and allopatric genotypes of I. vespertilionis in caves of Hungary. Parasit Vectors. 2014 Apr 27;7:202. doi: 10.1186/1756-3305-7-202.

Farkas R, Solymosi N, Takács N, Hornyák Á, Hornok S, Nachum-Biala Y, Baneth G. First molecular evidence of Hepatozoon canis infection in red foxes and golden jackals from Hungary. Parasit Vectors. 2014 Jul 2;7:303. doi: 10.1186/1756-3305-7-303.

Gyuranecz M, Sulyok K, Balla E, Mag T, Balazs A, Simor Z, Denes B, Hornok S, Bajnoczi P, Hornstra H, Pearson T, Keim P, Dan A. Q fever epidemic in Hungary, April to July 2013. Euro Surveill. 2014 Jul 31;19(30):20863. doi: 10.2807/1560-7917.es2014.19.30.20863.

Hornok S, Mester A, Takács N, Fernández de Mera IG, de la Fuente J, Farkas R. Re-emergence of bovine piroplasmosis in Hungary: has the etiological role of Babesia divergens been taken over by B. major and Theileria buffeli? Parasit Vectors. 2014 Sep 11;7:434. doi: 10.1186/1756-3305-7-434.

Hornok S, Abichu G, Meli ML, Tánczos B, Sulyok KM, Gyuranecz M, Gönczi E, Farkas R, Hofmann-Lehmann R. Influence of the biotope on the tick infestation of cattle and on the tick-borne pathogen repertoire of cattle ticks in Ethiopia. PLoS One. 2014 Sep 23;9(9):e106452. doi: 10.1371/journal.pone.0106452.

Sulyok KM, Hornok S, Abichu G, Erdélyi K, Gyuranecz M. Identification of novel Coxiella burnetii genotypes from Ethiopian ticks. PLoS One. 2014 Nov 25;9(11):e113213. doi: 10.1371/journal.pone.0113213.

Dumitrache MO, Paştiu AI, Kalmár Z, Mircean V, Sándor AD, Gherman CM, Peştean C, Mihalca AD, Cozma V. Northern white-breasted hedgehogs Erinaceus roumanicus as hosts for ticks infected with Borrelia burgdorferi sensu lato and Anaplasma phagocytophilum in Romania. Ticks Tick Borne Dis. 2013 Apr;4(3):214-7. doi: 10.1016/j.ttbdis.2012.11.010.

Farkas R, Tánczos B, Gyurkovszky M, Földvári G, Solymosi N, Edelhofer R, Hornok S. Serological and molecular detection of Theileria equi infection in horses in Hungary. Vet Parasitol. 2013 Feb 18;192(1-3):143-8. doi: 10.1016/j.vetpar.2012.09.035.

Hornok S, Csörgő T, de la Fuente J, Gyuranecz M, Privigyei C, Meli ML, Kreizinger Z, Gönczi E, Fernández de Mera IG, Hofmann-Lehmann R. Synanthropic birds associated with high prevalence of tick-borne rickettsiae and with the first detection of Rickettsia aeschlimannii in Hungary. Vector Borne Zoonotic Dis. 2013 Feb;13(2):77-83. doi: 10.1089/vbz.2012.1032.

Hornok S, Tánczos B, Fernández de Mera IG, de la Fuente J, Hofmann-Lehmann R, Farkas R. High prevalence of Hepatozoon-infection among shepherd dogs in a region considered to be free of Rhipicephalus sanguineus. Vet Parasitol. 2013 Sep 1;196(1-2):189-93. doi: 10.1016/j.vetpar.2013.02.009.

Hornok S, Dénes B, Meli ML, Tánczos B, Fekete L, Gyuranecz M, de la Fuente J, de Mera IG, Farkas R, Hofmann-Lehmann R. Non-pet dogs as sentinels and potential synanthropic reservoirs of tick-borne and zoonotic bacteria. Vet Microbiol. 2013 Dec 27;167(3-4):700-3. doi: 10.1016/j.vetmic.2013.08.011.

Gherman CM, Sándor AD, Kalmár Z, Marinov M, Mihalca AD. First report of Borrelia burgdorferi sensu lato in two threatened carnivores: the marbled polecat, Vormela peregusna and the European mink, Mustela lutreola (Mammalia: Mustelidae). BMC Vet Res. 2012 Aug 18;8:137. doi: 10.1186/1746-6148-8-137.

Dumitrache MO, Gherman CM, Cozma V, Mircean V, Györke A, Sándor AD, Mihalca AD. Hard ticks (Ixodidae) in Romania: surveillance, host associations, and possible risks for tick-borne diseases. Parasitol Res. 2012 May;110(5):2067-70. doi: 10.1007/s00436-011-2703-y.

Paștiu AI, Matei IA, Mihalca AD, D'Amico G, Dumitrache MO, Kalmár Z, Sándor AD, Lefkaditis M, Gherman CM, Cozma V. Zoonotic pathogens associated with Hyalomma aegyptium in endangered tortoises: evidence for host-switching behaviour in ticks? Parasit Vectors. 2012 Dec 28;5:301. doi: 10.1186/1756-3305-5-301.

Mihalca AD, Dumitrache MO, Magdaş C, Gherman CM, Domşa C, Mircean V, Ghira IV, Pocora V, Ionescu DT, Sikó Barabási S, Cozma V, Sándor AD. Synopsis of the hard ticks (Acari: Ixodidae) of Romania with update on host associations and geographical distribution. Exp Appl Acarol. 2012 Oct;58(2):183-206. doi: 10.1007/s10493-012-9566-5.

Hornok S, Horváth G, Jongejan F, Farkas R. Ixodid ticks on ruminants, with on-host initiated moulting (apolysis) of Ixodes, Haemaphysalis and Dermacentor larvae. Vet Parasitol. 2012 Jun 8;187(1-2):350-3. doi: 10.1016/j.vetpar.2011.12.012.

Hornok S, Kovács R, Meli ML, Gönczi E, Hofmann-Lehmann R, Kontschán J, Gyuranecz M, Dán A, Molnár V. First detection of bartonellae in a broad range of bat ectoparasites. Vet Microbiol. 2012 Oct 12;159(3-4):541-3. doi: 10.1016/j.vetmic.2012.04.003.

Hornok S, Horváth G. First report of adult Hyalomma marginatum rufipes (vector of Crimean-Congo haemorrhagic fever virus) on cattle under a continental climate in Hungary. Parasit Vectors. 2012 Aug 13;5:170. doi: 10.1186/1756-3305-5-170.

Demeter Z, Palade EA, Balogh E, Jakab C, Farkas R, Tánczos B, Hornok S. Postmortem small babesia-like morphology of Babesia canis - short communication. Acta Vet Hung. 2011 Dec;59(4):427-32. doi: 10.1556/AVet.2011.029.

Hornok S, Micsutka A, Fernández de Mera IG, Meli ML, Gönczi E, Tánczos B, Mangold AJ, Farkas R, Lutz H, Hofmann-Lehmann R, de la Fuente J. Fatal bovine anaplasmosis in a herd with new genotypes of Anaplasma marginale, Anaplasma ovis and concurrent haemoplasmosis. Res Vet Sci. 2012 Feb;92(1):30-5. doi: 10.1016/j.rvsc.2010.10.011.

Sikutová S, Hornok S, Hubálek Z, Dolezálková I, Juricová Z, Rudolf I. Serological survey of domestic animals for tick-borne encephalitis and Bhanja viruses in northeastern Hungary. Vet Microbiol. 2009 Mar 30;135(3-4):267-71. doi: 10.1016/j.vetmic.2008.09.082.

Boretti FS, Perreten A, Meli ML, Cattori V, Willi B, Wengi N, Hornok S, Honegger H, Hegglin D, Woelfel R, Reusch CE, Lutz H, Hofmann-Lehmann R. Molecular Investigations of Rickettsia helvetica infection in dogs, foxes, humans, and Ixodes ticks. Appl Environ Microbiol. 2009 May;75(10):3230-7. doi: 10.1128/AEM.00220-09.

Hornok S, Földvári G, Elek V, Naranjo V, Farkas R, de la Fuente J. Molecular identification of Anaplasma marginale and rickettsial endosymbionts in blood-sucking flies (Diptera: Tabanidae, Muscidae) and hard ticks (Acari: Ixodidae). Vet Parasitol. 2008 Jul 4;154(3-4):354-9. doi: 10.1016/j.vetpar.2008.03.019.

Hornok S, Elek V, de la Fuente J, Naranjo V, Farkas R, Majoros G, Földvári G. First serological and molecular evidence on the endemicity of Anaplasma ovis and A. marginale in Hungary. Vet Microbiol. 2007 Jun 21;122(3-4):316-22. doi: 10.1016/j.vetmic.2007.01.024.

Hornok S, Edelhofer R, Földvári G, Joachim A, Farkas R. Serological evidence for Babesia canis infection of horses and an endemic focus of B. caballi in Hungary. Acta Vet Hung. 2007 Dec;55(4):491-500. doi: 10.1556/AVet.55.2007.4.8.

Hornok S, Edelhofer R, Farkas R. Seroprevalence of canine babesiosis in Hungary suggesting breed predisposition. Parasitol Res. 2006 Nov;99(6):638-42. doi: 10.1007/s00436-006-0218-8.

Hornok S, Edelhofer R, Szotáczky I, Hajtós I. Babesia divergens becoming extinct in cattle of Northeast Hungary: new data on the past and present situation. Acta Vet Hung. 2006 Dec;54(4):493-501. doi: 10.1556/AVet.54.2006.4.7.

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